The role of the player's body is really interesting. It's always interesting to see where different players, especially better players, tune. Usually the better the player is, the further in they tune (relative to others, on the same mouthpiece), but not always. Thanks for that thorough answer.
RE toneholes, I also ran some experiments re...what do I call him..."the Phantom of the Benade Opera?"
Anyway, I forget how I went about it, but there was a recommendation, widely distributed through internet forum posts about altering the volume of toneholes -- encouraging it, I think -- specificially about palm key toneholes. This was at least 10 years ago. I forget, now, exactly what the suggestion was, but I discarded it after running some experiments of it on my own. The idea was correct, I remember: whatever altering the volume of the tonehole was supposed to do, it at least somewhat did, but it also threw the intonation of several other notes behind it, and the whole experience of playing the horn out of whack (this was done in a reversible experiment -- nobody inherited a horn that suffered permanent damage from that dead end).